Mammalia - Primates - Cercopithecidae

Parent taxon: Colobinae according to M. Osterholz et al. 2008

Sister taxa: Colobus, Dolichopithecus, Mesopithecus, Nasalis, Presbytis, Procolobus, Rhinopithecus, Semnopithecus, Simias, Trachypithecus

Subtaxa: Pygathrix nemaeus, Pygathrix nigripes

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Ecology: arboreal frugivore-folivore