Full reference: J. A. Clarke, D. T. Ksepka, M. Stucchi, M. Urbina, N. Giannini, S. Bertelli, Y. Narvaez and C. A. Boyd. 2007. Paleogene equatorial penguins challenge the proposed relationship between biogeography, diversity and Cenozoic climate change. Proceedings of the Academy of Natural Sciences 104(28):11545-11550
Sister taxa: none
Type specimen: MUSM 889, a partial skeleton. Its type locality is Quebrada Perdida, which is in a Lutetian marginal marine sandstone in the Paracas Formation of Peru.
Ecology: ground dwelling carnivore