Coudoulous I (Faunal Unit III)
Basic information
Sample name: Coudoulous I (Faunal Unit III)

Reference: P. Fernandez, M. Jeannet, J. Jaubert, and J.-P. Brugal. 2021. Paleoecological first results on the Middle Pleistocene sequence of Coudoulous I (Quercy, Lot, France). Alpine and Mediterranean Quaternary 34(1):89-107 [ER 3857]
Geography
Country: France

Coordinate: 44° 28' 20" N, 2° 40' 50" E
Latlng basis: stated in text

Time interval: Middle Pleistocene

Max Ma: 0.229

Min Ma: 0.1613

Age basis: other

Geography comments: "in the Quercy area... at the junction of the river LOt and its tributary, the Célé"
the entire faunal sequence is bracketed by an ESR/U-series date of 161.3 +/- 14 kyr at the top and two U/Th dates of 229 + 78 - 42 and 202 + 78 - 39 kyr at the base

Environment
Lithology: claystone

Taphonomic context: cave, human accumulation, pitfall trap

Archaeology: stone tools

Habitat comments: a pitfall trap with fossils from "clays, sands, angular limestone rocks"
there is a "very rich archaeological level" immediately above the fossiliferous sequence that dates to the "Early Middle Paleolithic" and there are "scarce and dispersed artefacts of the same age" in the fossiliferous levels
the site "was used for marginal scavenging by Lower Paleolithic hominid groups and as a kill-butchery site during the EMP", so it is at least partially a human accumulation
from stratigraphic units 6a, 7a, 7a', and 7b
there is apparently no sandstone in this interval, so the fossils are presumably from clays

Methods
Life forms: bats, carnivores, rodents, ungulates, other small mammals, lizards, snakes, frogs, salamanders

Sampling methods: quarry, screenwash

Sample size: 3673 specimens

Years: 1994 - 2003

Net or trap nights: 0

Basal area status: not applicable

Sampling comments: there was a "rescue excavation" in 1978 - 1980, but the main excavation was between 1994 and 2003
the high abundance of micromammals is taken to mean that screening was used

Metadata
Sample no: 4224

Contributor no: John Alroy

Enterer: John Alroy

Created: 2023-04-14 17:42:54

Modified: 2023-04-14 07:49:18

Abundance distribution
Each square represents a species. Square sizes are proportional to counts. Values are logged.
Statistics
73 species
16 singletons
total count 3673
geometric series index: 111.7
Fisher's α: 12.911
geometric series k: 0.909
Hurlbert's PIE: 0.8835
Shannon's H: 2.7809
Good's u: 0.9956
Register
Microtus arvalis 959
Microtus agrestis 284
Stenocranius gregalis 234
"Microtus gregalis"
Alexandromys oeconomus 39
"Microtus oeconomus"
Microtus brecciensis 77
Microtus multiplex 54
Microtus gerbii 39
"Microtus pyrenaicus"
Microtus subterraneus 51
Myodes glareolus 82
"Clethrionomys glareolus"
Lemmus lemmus 1
Pliomys lenki 26
Arvicola amphibius 4
"Arvicola terrestris"
Arvicola cantiana 33
Allocricetus bursae 146
Apodemus sylvaticus 599
Apodemus flavicollis 6
Apodemus uralensis 47
Eliomys quercinus 202
Muscardinus avellanarius 8
Erinaceus davidi 2
Talpa europaea 10
Talpa minor 1
Sorex araneus 7
Sorex minutus 8
Sorex minutissimus 6
Crocidura russula 5
Oryctolagus cuniculus 82.5 kg
Lepus europaeus 33.4 kg
Lepus timidus 2
Ochotona pusilla 9
Myotis myotis 125.7 g
Myotis capaccinii 110.0 g
Pipistrellus pipistrellus 14.4 g
Barbastella barbastellus 28.0 g
Plecotus auritus 17.6 g
Salamandra salamandra 2
Bombina variegata 1
Bufo bufo 2
Bufo calamita 7
Bufo viridis 3
Hyla arborea 1
Rana dalmatina 6
Rana temporaria 3
Lacerta viridis 4
Podarcis cf. muralis 2
Anguis fragilis 7
Coronella austriaca 2
Coronella girondica 1
Malpolon monspessulanus 1
Natrix maura 2
Natrix natrix 1
Vipera berus 3
Cervus elaphus (red deer)88
Dama dama (European fallow deer)4
Capreolus capreolus (roe deer)17
Bison priscus (bison)51
compared to subspecies mediator
Equus cf. mosbachensis (horse)30
Hemitragus cedrensis (tahr)48
Palaeoloxodon antiquus (elephant)3
Stephanorhinus cf. hemitoechus (narrow-nosed rhinoceros)7
Rupicapra sp. 1
Sus scrofa (pig)1
Canis lupus lupus (gray wolf)21443.0 kg
Cuon alpinus (dhole)5
Vulpes vulpes (red fox)9
Felis silvestris (European wildcat)14.5 kg
Lynx lynx (Eurasian lynx)7
Panthera spelaea (cave lion)56
Ursus arctos (brown bear)56
Panthera pardus (leopard)1
Mustela nivalis (least weasel)6061.7 g
Mustela putorius (European polecat)1
Meles meles (European badger)711.0 kg