Coudoulous I (Faunal Units IV and V)
Basic information
Sample name: Coudoulous I (Faunal Units IV and V)

Reference: P. Fernandez, M. Jeannet, J. Jaubert, and J.-P. Brugal. 2021. Paleoecological first results on the Middle Pleistocene sequence of Coudoulous I (Quercy, Lot, France). Alpine and Mediterranean Quaternary 34(1):89-107 [ER 3857]
Geography
Country: France

Coordinate: 44° 28' 20" N, 2° 40' 50" E
Latlng basis: stated in text

Time interval: Middle Pleistocene

Max Ma: 0.229

Min Ma: 0.1613

Age basis: other

Geography comments: "in the Quercy area... at the junction of the river LOt and its tributary, the Célé"
the entire faunal sequence is bracketed by an ESR/U-series date of 161.3 +/- 14 kyr at the top and two U/Th dates of 229 + 78 - 42 and 202 + 78 - 39 kyr at the base

Environment
Lithology: claystone

Taphonomic context: cave, human accumulation, pitfall trap

Archaeology: stone tools

Habitat comments: a pitfall trap with fossils from "clays, sands, angular limestone rocks"
there is a "very rich archaeological level" immediately above the fossiliferous sequence that dates to the "Early Middle Paleolithic" and there are "scarce and dispersed artefacts of the same age" in the fossiliferous levels
the site "was used for marginal scavenging by Lower Paleolithic hominid groups and as a kill-butchery site during the EMP", so it is at least partially a human accumulation
from stratigraphic units 7c, 7d, 7x, 7e, and 7f
there is apparently no sandstone in this interval, so the fossils are presumably from clays

Methods
Life forms: bats, carnivores, rodents, ungulates, other small mammals, lizards, snakes, frogs, salamanders

Sampling methods: quarry, screenwash

Sample size: 3226 specimens

Years: 1994 - 2003

Net or trap nights: 0

Basal area status: not applicable

Sampling comments: there was a "rescue excavation" in 1978 - 1980, but the main excavation was between 1994 and 2003
the high abundance of micromammals is taken to mean that screening was used

Metadata
Sample no: 4225

Contributor no: John Alroy

Enterer: John Alroy

Created: 2023-04-14 17:52:54

Modified: 2023-04-14 07:52:54

Abundance distribution
Each square represents a species. Square sizes are proportional to counts. Values are logged.
Statistics
96 species
18 singletons
total count 3226
geometric series index: 148.8
Fisher's α: 18.599
geometric series k: 0.9357
Hurlbert's PIE: 0.9221
Shannon's H: 3.0879
Good's u: 0.9944
Register
Microtus arvalis 553
Microtus agrestis 74
Stenocranius gregalis 265
"Microtus gregalis"
Alexandromys oeconomus 43
"Microtus oeconomus"
Chionomys nivalis 3
Microtus brecciensis 68
Microtus multiplex 26
Microtus gerbii 9
"Microtus pyrenaicus"
Microtus subterraneus 30
Myodes glareolus 28
"Clethrionomys glareolus"
Lemmus lemmus 1
Dicrostonyx torquatus 252.0 g
Pliomys lenki 22
Pliomys chalinei 1
Arvicola amphibius 2
"Arvicola terrestris"
Arvicola cantiana 15
Allocricetus bursae 165
Apodemus sylvaticus 293
Apodemus flavicollis 7
Apodemus uralensis 21
Eliomys quercinus 400
Muscardinus avellanarius 4
Sciurus vulgaris 1311.0 g
Erinaceus europaeus 3
Erinaceus davidi 4
Talpa europaea 9
Talpa minor 1
Sorex araneus 6
Sorex minutus 7
Sorex minutissimus 1
Crocidura russula 6
Crocidura leucodon 1
Crocidura suaveolens 1
Castor cf. fiber 1
Oryctolagus cuniculus 62.5 kg
Lepus europaeus 83.4 kg
Lepus timidus 5
Ochotona pusilla 7
Rhinolophus ferrumequinum 120.0 g
Rhinolophus hipposideros 15.3 g
Myotis myotis 325.7 g
Myotis blythii 224.6 g
Myotis capaccinii 110.0 g
Myotis mystacinus 25.5 g
Myotis bechsteinii 18.5 g
Vespertilio murinus 111.9 g
Eptesicus serotinus 122.6 g
Nyctalus noctula 129.2 g
Pipistrellus pipistrellus 24.4 g
Barbastella barbastellus 38.0 g
Plecotus auritus 57.6 g
Plecotus austriacus 38.3 g
Salamandra salamandra 4
Pelobates fuscus 2
Bufo bufo 6
Bufo calamita 6
Hyla arborea 2
Rana dalmatina 4
Rana temporaria 3
Lacerta agilis 2
Lacerta viridis 8
Podarcis cf. muralis 3
Chalcides chalcides 2
Anguis fragilis 8
Coluber viridiflavus 3
Coronella austriaca 6
Coronella girondica 6
Elaphe longissima 2
Rhinechis scalaris 1
Malpolon monspessulanus 2
Natrix maura 2
Natrix natrix 6
Vipera aspis 3
Vipera berus 5
Vipera ursinii 3
Cervus elaphus (red deer)114
Dama dama (European fallow deer)14
Capreolus capreolus (roe deer)15
Bison priscus (bison)99
compared to subspecies mediator
Equus cf. mosbachensis (horse)39
Hemitragus cedrensis (tahr)50
Palaeoloxodon antiquus (elephant)23
Stephanorhinus cf. hemitoechus (narrow-nosed rhinoceros)8
Sus scrofa (pig)1
Canis lupus lupus (gray wolf)28943.0 kg
Cuon alpinus (dhole)16
Vulpes vulpes (red fox)9
Felis silvestris (European wildcat)24.5 kg
Lynx lynx (Eurasian lynx)2
Panthera spelaea (cave lion)96
Ursus arctos (brown bear)117
Panthera pardus (leopard)2
Mustela erminea (stoat)1
Mustela nivalis (least weasel)10261.7 g
Mustela putorius (European polecat)4
Meles meles (European badger)811.0 kg