Latitudinal range: 36.8° to 72.8°

Habitats: Mediterranean woodland (1), boreal forest/taiga (1)

Found in 9 samples

Canada: Churchill Falls, Charlie Lake Cave (Zone IV) (Holocene), Learmonth Site (Holocene), McNichol Creek (house depression) (Holocene), McNichol Creek (midden) (Holocene)

France: Piémanson Beach

Spain: Cueva de Nerja (Tardiglacial) (Late Pleistocene), Cueva de Nerja (Early Holocene) (Holocene)

United States: McIntosh (Holocene)

See also Larus argentatus, Larus canus, Larus fuscus, Larus hyperboreus, Larus marinus, Larus melanocephalus, Larus michahellis, Larus sp.