Habitats: temperate broadleaf/mixed forest (2), Mediterranean woodland (2)
Found in 12 samples
France: Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Coudoulous I (Faunal Unit VII) (Middle Pleistocene)
Greece: Lake Prespa National Park
Italy: Poggi di Prata, Cava Muracci (Late Pleistocene)
Montenegro: Great Ulcinj Beach
Poland: Zabia Cave (Layer 21) (Early Pleistocene), Zabia Cave (Layer 19) (Early Pleistocene)
Turkey: Madran Mountain
Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)
See also Natrix, Natrix hillmani, Natrix longivertebrata, Natrix maura, Natrix merkurensis, Natrix sansaniensis, Natrix sipedon, Natrix sp., Natrix sp. 1, Natrix sp. 2, Natrix tessellata