Latitudinal range: 38.5° to 43.1°

Habitats: temperate broadleaf/mixed forest (2), Mediterranean woodland (1)

Found in 6 samples

Greece: Lake Prespa National Park

Italy: Poggi di Prata

Montenegro: Great Ulcinj Beach

Portugal: Gruta de Figueira Brava (Area C) (Late Pleistocene)

Spain: Bolomor Cave (Level XI) (Middle Pleistocene), Bolomor Cave (Level IV) (Middle Pleistocene)

See also Testudo, Testudo brevitesta, Testudo burgenlandica, Testudo graeca, Testudo marmorum, Testudo s.l., Testudo sp.