Latitudinal range: 41.9° to 50.6°

Habitats: temperate broadleaf/mixed forest (1), Mediterranean woodland (1)

Found in 7 samples

Italy: Poggi di Prata

Montenegro: Great Ulcinj Beach

Poland: Borsuka Cave (Layer II) (Holocene), Zabia Cave (Layer 19) (Early Pleistocene), Zabia Cave (Layer 17) (Early Pleistocene), Zabia Cave (Layer 15) (Early Pleistocene)

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

See also Zamenis, Zamenis scalaris