Abundance: 0.02 to 27.27% (median 0.55%)

Latitudinal range: 32.2° to 51.4°

Habitats: (21), Mediterranean woodland (1), temperate broadleaf/mixed forest (3)

Found in 25 samples

Czech Republic: Merkur−North (Early Miocene)

France: Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene)

Germany: Hambach 6C (Middle Miocene), Griesbeckerzell 1a (Middle Miocene), Griesbeckerzell 1b (Middle Miocene)

Greece: Lake Prespa National Park

Iran: Wezmeh Cave (Late Pleistocene)

Italy: Poggi di Prata, Monte Argentario (Early Pleistocene), Monticino Quarry (Late Miocene)

Jordan: Dibbeen Nature Reserve

Kazakhstan: Baikadam (Middle Miocene)

Montenegro: Great Ulcinj Beach

Poland: Zabia Cave (Layer 21) (Early Pleistocene), Zabia Cave (Layer 17) (Early Pleistocene), Zabia Cave (Layer 15) (Early Pleistocene)

Romania: Subpiatra 2/1R (Middle Miocene)

Spain: Caló den Rafelino (Early Pliocene), Puerto de la Cadena (Early Pliocene)

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

See also Vipera ammodytes, Vipera aspis, Vipera berus, Vipera lebetina, Vipera natiensis, Vipera palaestinae, Vipera s.l., Vipera sp., Vipera sp. 2, Vipera ursinii