Abundance: 0.02 to 27.27% (median 1.38%)

Latitudinal range: 37.9° to 51.4°

Habitats: (11) only

Found in 11 samples

Czech Republic: Merkur−North (Early Miocene)

France: Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene)

Germany: Hambach 6C (Middle Miocene), Griesbeckerzell 1a (Middle Miocene), Griesbeckerzell 1b (Middle Miocene)

Italy: Monte Argentario (Early Pleistocene)

Kazakhstan: Baikadam (Middle Miocene)

Romania: Subpiatra 2/1R (Middle Miocene)

Spain: Puerto de la Cadena (Early Pliocene)

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

See also Vipera, Vipera ammodytes, Vipera aspis, Vipera berus, Vipera lebetina, Vipera natiensis, Vipera palaestinae, Vipera s.l., Vipera sp. 2, Vipera ursinii